Dec 05, 2025  
2025-2026 Academic Catalog 
    
2025-2026 Academic Catalog

Biology, B.A.


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Biology 

Member of the Division of Science

Chair:

Joshua Sandquist

Faculty:

Charvann Bailey
David Campbell
Idelle Cooper
Benjamin DeRidder
Caroline Dong
Vince Eckhart
Leslie Gregg-Jolly (senior faculty status)
Shannon Hinsa

Kathy Jacobson
Peter Jacobson
Pascal Lafontant
Clark Lindgren
Vida Praitis
Elizabeth Queathem


 

Our biology curriculum is among the nation’s most innovative. From their very first biology class, Grinnell students learn how to think and act like biologists, actively investigating and interpreting the unity and diversity of life. Acknowledging that biology intersects with diverse disciplines and with societal issues, we encourage students to gain knowledge of other sciences and mathematics, to explore interdisciplinary issues, to participate in off-campus study, and to carry out independent research (in faculty research labs, at Grinnell’s Conard Environmental Research Area, or off-campus). In this way Grinnell biology students prepare for advanced study or careers in education, health professions, research, environmental science, biotechnology, conservation, and other fields. Our Department’s mission and its associated learning goals are accomplished through a curriculum that integrates teaching and research at all levels.

Mission Statement: Students studying biology at Grinnell College practice the methods of biological inquiry to uncover the wonder of life, from molecules to ecosystems, and from the foundations to the frontiers of the life sciences. Students are well-placed to use this knowledge to understand the relevance of biology to other disciplines, society, and emerging challenges.

Learning Outcomes. To accomplish our mission, we articualte the following learning outcomes for students who progress through our curriculum:

  1. Students conduct biological inquiry proficiently, as exhibited by:

a. designing and conducting investigations to ask questions about living systems.
b. evaluating data and other forms of evidence pursuant to raising a novel question or hypothesis.
c. effectively and responsibly communicating biological information and ideas.
d. identifying, evaluating, and properly integrating knowledge from the scientific literature.

2. Students apply key biological concepts to all levels of biological organization: 

a. relating physical and chemical characteristics of living things to biological function.
b. analyzing how sub-organismal components independently and collectively support organismal function.
c. explaining how organisms survive in, reproduce in, interact with, and evolve in their environment.
d. applying evolutionary theory to understand the phylogenetic relationships of organisms.
e. applying elements of organismal structure to organismal function in populations, communities, and ecosystems.

3. Students connect biology to other academic disciplines and society, as exhibited by:

a. applying biological principles to problems that affect human society.
b. evaluating how societal and cultural practices and beliefs influence the practice of biological research.

Curriculum. Biology investigates the processes of life at the molecular, cellular, organismal, population, and ecosystem levels. The study of biology can help students of all majors and interests become scientifically literate, and our curriculum centers on the practice of biological science. Students in our introductory course engage in authentic research by making original observations, designing and executing investigations that address important biological questions, analyzing and interpreting scientific data, and communicating their findings. Intermediate-level courses expose students to the central concepts and diverse approaches that underpin the full range of biological organization, from molecules to ecosystems. Advanced courses provide in-depth opportunities to explore a broad spectrum of biology sub-disciplines, enriching students’ ability to read and write scientific literature, to think independently while working collaboratively, and to prepare for future work.

We recommend that students considering a biology (or biological chemistry) major take both BIO 150 - Introduction to Biological Inquiry and CHM 129 - General Chemistry in their first year, in either order. We also recommend that first-year students take calculus ( MAT 123 - Functions and Differential Calculus – MAT 124 - Functions and Integral Calculus or MAT 131 - Calculus I) or applied statistics (STA 209). In the first semester of their second year, students should take both BIO 251 - Molecules, Cells, and Organisms and its co-requisite, CHM 221 - Organic Chemistry I. In the second semester, students continuing as biology majors should enroll in BIO 252 - Organisms, Evolution, and Ecology. In the third and fourth years, careful planning with faculty advisers will help students schedule a set of appropriate advanced electives, also considering options for independent research in biology and off-campus study.

Excellent laboratory and field facilities support the biology program. Biology Department space in the Noyce Science Center includes classrooms, teaching labs, faculty research labs, a greenhouse, a herbarium, and numerous support rooms designed explicitly to facilitate the research-intensive curriculum. Instrumentation available for courses and research includes confocal and wide-field fluorescence microscopes, high-speed and ultra centrifuges, a liquid scintillation spectrometer, scanning UV-visible spectrophotometers and plate readers, luminometer, electrophysiological suites and tissue bath systems, C/N, TOC and FIA autoanalyzers, mammalian cell-culture equipment, an Odyssey CLx infrared imaging system, and a quantitative, real-time PCR system, as well as standard equipment for molecular biology. The department also manages the Conard Environmental Research Area (CERA), a 148-hectare (365-acre) biological field station 11 miles from campus, which includes oak woodland, savanna, tallgrass prairie, diverse aquatic habitats, and several long-term experimental areas. The Environmental Education Center at CERA supports teaching and research in biology, the arts, and other subjects with excellent classroom, collection, and greenhouse facilities.


Major Requirements: a minimum of 32 credits


 

Additional Courses: 20 credits


  • 20 credits of 200-level or higher elective courses in Biology or BCM 262  
  • At least 12 of the credits must be at the 300-level or higher
  • At least 12 of the credits must include a lab component
  • At least 12 of the credits must be Grinnell courses

Additional Information


CHM 222 , PHY 131 , PHY 132 , and STA 209  are recommended.

Not more than 5 of the 20 credits may be taken from BIO 297, BIO 299, BIO 397, BIO 399, BIO 499, SCI 300 (see additional information in Individual Study, Internships, and Practica ) or independent study done elsewhere. 

With prior approval, a maximum of four credits of advanced work in a related field may be applied toward the major. The following courses are on the approved list: ANT 221 , NRS 250 PSY 336 .

 

Honors


To be considered for honors in biology, graduating seniors, in addition to meeting the College’s general requirements for honors, must conduct an independent research project (either at Grinnell or elsewhere) and share their findings with fellow biologists in a departmental seminar. The award of honors is not based solely on grades and achievement in the classroom or lab. It signifies, in addition, an underlying commitment to the discipline as evidenced by participation in departmental affairs and activities (e.g., acting as a teaching assistant or mentor, or serving on the SEPC), including regular attendance at departmental seminars.